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Can A Turtle Run . This speed varies from one species to another. When startled, they can run very quickly. Back 4 Blood trailer details the game's roguelike card system that from www.vg247.com Apparently, the greatest speed that a tortoise has ever reached is a whopping 0.63 mph. Turtles can run at speeds of up to 3 miles per hour. Turtles can run, not very fast, but they can run!